:root {
  --ink: #14221e;
  --muted: #5c6b66;
  --line: #dfe8e4;
  --paper: #ffffff;
  --wash: #f4f8f6;
  --green: #183f35;
  --green-2: #2f6b58;
  --blue: #315c85;
  --burgundy: #7a3e4a;
  --gold: #c8a84f;
  --shadow: 0 18px 44px rgba(16, 34, 29, 0.1);
  --max: 1180px;
}

* {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
}

body {
  margin: 0;
  color: var(--ink);
  background: var(--wash);
  font-family:
    Inter, ui-sans-serif, system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  line-height: 1.6;
}

a {
  color: inherit;
  text-decoration: none;
}

button,
a {
  -webkit-tap-highlight-color: transparent;
}

.site-header {
  position: sticky;
  top: 0;
  z-index: 20;
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 24px;
  min-height: 72px;
  padding: 12px max(22px, calc((100vw - var(--max)) / 2));
  border-bottom: 1px solid rgba(223, 232, 228, 0.74);
  background: rgba(255, 255, 255, 0.9);
  backdrop-filter: blur(18px);
}

.brand {
  display: inline-flex;
  align-items: center;
  gap: 12px;
  min-width: 0;
}

.brand-logo {
  display: block;
  width: 116px;
  height: 50px;
  object-fit: contain;
}

.brand-logo[hidden] {
  display: none;
}

.brand-mark {
  display: grid;
  place-items: center;
  width: 42px;
  height: 42px;
  border: 1px solid rgba(47, 107, 88, 0.35);
  border-radius: 8px;
  color: var(--paper);
  background: linear-gradient(135deg, var(--green), var(--blue));
  font-size: 0.9rem;
  font-weight: 800;
}

.brand-mark[hidden] {
  display: none;
}

.brand strong,
.brand small {
  display: block;
  white-space: nowrap;
}

.brand strong {
  font-size: 0.96rem;
  line-height: 1.15;
}

.brand small {
  color: var(--muted);
  font-size: 0.78rem;
}

.site-nav {
  display: flex;
  align-items: center;
  gap: 4px;
}

.site-nav a {
  padding: 10px 12px;
  border-radius: 6px;
  color: #31423d;
  font-size: 0.95rem;
}

.site-nav a:hover {
  background: rgba(47, 107, 88, 0.1);
}

.nav-toggle {
  display: none;
  width: 42px;
  height: 42px;
  place-items: center;
  border: 1px solid var(--line);
  border-radius: 6px;
  color: var(--green);
  background: var(--paper);
  cursor: pointer;
}

.nav-toggle svg,
.icon-button svg,
.button svg,
.contact-link svg,
.research-card svg {
  width: 18px;
  height: 18px;
  stroke-width: 1.9;
}

.hero {
  display: flex;
  align-items: center;
  min-height: 84vh;
  padding: 108px max(22px, calc((100vw - var(--max)) / 2)) 92px;
  color: var(--paper);
  background:
    linear-gradient(90deg, rgba(15, 37, 32, 0.96) 0%, rgba(15, 37, 32, 0.86) 34%, rgba(15, 37, 32, 0.24) 76%),
    url("assets/hero-heor-lab.png") center right / cover no-repeat;
}

.hero-copy {
  width: min(690px, 100%);
}

.eyebrow,
.section-kicker {
  margin: 0 0 12px;
  color: var(--gold);
  font-size: 0.78rem;
  font-weight: 800;
  text-transform: uppercase;
}

.hero h1,
.section-heading h2,
.section-grid h2,
.contact-inner h2 {
  margin: 0;
  line-height: 1.05;
  font-family: Georgia, "Times New Roman", serif;
  font-weight: 700;
}

.hero h1 {
  max-width: 760px;
  font-size: 4rem;
}

.hero-lede {
  max-width: 650px;
  margin: 24px 0 0;
  color: rgba(255, 255, 255, 0.9);
  font-size: 1.2rem;
}

.hero-actions {
  display: flex;
  flex-wrap: wrap;
  gap: 12px;
  margin-top: 34px;
}

.button,
.icon-button,
.contact-link {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  gap: 8px;
  min-height: 44px;
  border-radius: 6px;
  font-weight: 800;
  transition:
    transform 160ms ease,
    border-color 160ms ease,
    background 160ms ease;
}

.button {
  padding: 0 18px;
  border: 1px solid transparent;
}

.button:hover,
.icon-button:hover,
.contact-link:hover {
  transform: translateY(-1px);
}

.button.primary {
  color: var(--green);
  background: var(--paper);
}

.button.secondary {
  color: var(--paper);
  border-color: rgba(255, 255, 255, 0.42);
  background: rgba(255, 255, 255, 0.1);
}

.button.compact {
  min-height: 40px;
  padding: 0 12px;
  color: var(--green);
  border-color: var(--line);
  background: var(--paper);
  font-size: 0.9rem;
}

.intro-section,
.research-section,
.publications-section,
.people-section,
.contact-section {
  padding: 86px max(22px, calc((100vw - var(--max)) / 2));
}

.intro-section,
.publications-section {
  background: var(--paper);
}

.section-grid {
  display: grid;
  grid-template-columns: minmax(220px, 0.75fr) minmax(0, 1.25fr);
  gap: 64px;
  align-items: start;
}

.section-grid h2,
.section-heading h2,
.contact-inner h2 {
  font-size: 2.55rem;
}

.intro-copy {
  display: grid;
  gap: 18px;
  color: #2d3c38;
  font-size: 1.06rem;
}

.intro-copy p {
  margin: 0;
}

.section-heading {
  display: flex;
  align-items: end;
  justify-content: space-between;
  gap: 32px;
  max-width: var(--max);
  margin: 0 auto 34px;
}

.research-grid {
  display: grid;
  grid-template-columns: repeat(4, minmax(0, 1fr));
  gap: 16px;
  max-width: var(--max);
  margin: 0 auto;
}

.research-card,
.publication-card,
.pi-card,
.team-notes article,
.member-card {
  border: 1px solid var(--line);
  border-radius: 8px;
  background: var(--paper);
}

.research-card {
  min-height: 270px;
  padding: 24px;
  box-shadow: var(--shadow);
}

.research-card svg {
  width: 30px;
  height: 30px;
  color: var(--green-2);
}

.research-card h3,
.publication-card h3,
.pi-card h3,
.team-notes h3,
.member-card h3 {
  margin: 18px 0 10px;
  line-height: 1.25;
}

.research-card p,
.team-notes p,
.pi-card p,
.member-card p {
  margin: 0;
  color: var(--muted);
}

.method-band {
  overflow: hidden;
  padding: 28px 0;
  color: var(--paper);
  background: linear-gradient(90deg, var(--green), var(--blue), var(--burgundy));
}

.method-track {
  display: flex;
  flex-wrap: wrap;
  justify-content: center;
  gap: 10px;
  width: min(var(--max), calc(100% - 44px));
  margin: 0 auto;
}

.method-track span {
  padding: 8px 12px;
  border: 1px solid rgba(255, 255, 255, 0.25);
  border-radius: 6px;
  background: rgba(255, 255, 255, 0.08);
  font-size: 0.9rem;
  font-weight: 700;
}

.publications-heading {
  align-items: center;
}

.publication-actions {
  display: flex;
  flex-wrap: wrap;
  justify-content: flex-end;
  gap: 8px;
}

.icon-button {
  width: 40px;
  height: 40px;
  border: 1px solid var(--line);
  color: var(--green);
  background: var(--paper);
  cursor: pointer;
}

.publication-meta {
  display: flex;
  flex-wrap: wrap;
  justify-content: space-between;
  gap: 10px;
  max-width: var(--max);
  margin: 0 auto 18px;
  color: var(--muted);
  font-size: 0.9rem;
}

.publication-toolbar {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 14px;
  max-width: var(--max);
  margin: 0 auto 18px;
}

.publication-tabs {
  display: inline-grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 4px;
  padding: 4px;
  border: 1px solid var(--line);
  border-radius: 8px;
  background: rgba(255, 255, 255, 0.74);
  box-shadow: 0 10px 28px rgba(16, 34, 29, 0.06);
}

.publication-tab {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  gap: 8px;
  min-height: 36px;
  padding: 0 14px;
  border: 0;
  border-radius: 6px;
  color: var(--muted);
  background: transparent;
  font: inherit;
  font-size: 0.88rem;
  font-weight: 800;
  cursor: pointer;
}

.publication-tab.is-active {
  color: var(--paper);
  background: linear-gradient(135deg, var(--green), var(--blue));
  box-shadow: 0 8px 18px rgba(24, 63, 53, 0.18);
}

.publication-tab span {
  min-width: 22px;
  padding: 1px 7px;
  border-radius: 999px;
  color: inherit;
  background: rgba(255, 255, 255, 0.18);
  font-size: 0.76rem;
}

.publication-summary {
  margin: 0;
  color: var(--muted);
  font-size: 0.88rem;
  font-weight: 700;
}

.publication-list {
  display: grid;
  gap: 12px;
  max-width: var(--max);
  margin: 0 auto;
}

.publication-list.is-archive {
  gap: 0;
}

.publication-card {
  padding: 20px 22px;
}

.publication-card h3 {
  margin-top: 0;
  font-size: 1.05rem;
}

.publication-card .source-line {
  margin: 0 0 8px;
  color: var(--green-2);
  font-size: 0.78rem;
  font-weight: 800;
  text-transform: uppercase;
}

.publication-card h3 a {
  text-decoration: underline;
  text-decoration-color: rgba(47, 107, 88, 0.34);
  text-underline-offset: 3px;
}

.publication-card p {
  margin: 8px 0 0;
  color: var(--muted);
}

.publication-card .journal-line {
  display: flex;
  flex-wrap: wrap;
  gap: 8px;
  color: #31423d;
  font-size: 0.92rem;
  font-weight: 800;
}

.publication-card .doi {
  color: var(--blue);
  font-size: 0.9rem;
}

.publication-archive {
  display: grid;
  gap: 18px;
}

.publication-year-group {
  display: grid;
  grid-template-columns: 92px minmax(0, 1fr);
  gap: 18px;
  align-items: start;
}

.publication-year {
  position: sticky;
  top: 92px;
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 38px;
  border: 1px solid rgba(47, 107, 88, 0.22);
  border-radius: 8px;
  color: var(--green);
  background: rgba(255, 255, 255, 0.78);
  font-family: Georgia, "Times New Roman", serif;
  font-size: 1.1rem;
  font-weight: 700;
}

.publication-year-items {
  position: relative;
  display: grid;
  gap: 10px;
  padding-left: 18px;
}

.publication-year-items::before {
  position: absolute;
  top: 10px;
  bottom: 10px;
  left: 0;
  width: 1px;
  background: linear-gradient(var(--green-2), rgba(49, 92, 133, 0.22));
  content: "";
}

.publication-row {
  position: relative;
  display: grid;
  grid-template-columns: minmax(0, 1fr) auto;
  gap: 16px;
  align-items: start;
  padding: 17px 18px;
  border: 1px solid rgba(223, 232, 228, 0.9);
  border-radius: 8px;
  background: rgba(255, 255, 255, 0.82);
  box-shadow: 0 12px 34px rgba(16, 34, 29, 0.06);
}

.publication-row::before {
  position: absolute;
  top: 24px;
  left: -23px;
  width: 9px;
  height: 9px;
  border: 2px solid var(--paper);
  border-radius: 50%;
  background: var(--green-2);
  box-shadow: 0 0 0 2px rgba(47, 107, 88, 0.22);
  content: "";
}

.publication-row h3 {
  margin: 0;
  font-size: 1rem;
  line-height: 1.35;
}

.publication-row h3 a {
  text-decoration: underline;
  text-decoration-color: rgba(47, 107, 88, 0.34);
  text-underline-offset: 3px;
}

.publication-row p {
  margin: 7px 0 0;
  color: var(--muted);
}

.publication-row .source-line {
  margin-top: 0;
  color: var(--green-2);
  font-size: 0.72rem;
  font-weight: 800;
  text-transform: uppercase;
}

.publication-row .journal-line {
  display: flex;
  flex-wrap: wrap;
  gap: 8px;
  color: #31423d;
  font-size: 0.9rem;
  font-weight: 800;
}

.publication-doi-chip {
  align-self: start;
  padding: 5px 9px;
  border: 1px solid rgba(49, 92, 133, 0.2);
  border-radius: 999px;
  color: var(--blue);
  background: rgba(49, 92, 133, 0.08);
  font-size: 0.72rem;
  font-weight: 800;
}

.skeleton {
  display: grid;
  gap: 10px;
}

.skeleton div {
  height: 14px;
  border-radius: 6px;
  background: linear-gradient(90deg, #edf3f0, #f8faf9, #edf3f0);
}

.skeleton div:first-child {
  width: 72%;
}

.skeleton div:nth-child(2) {
  width: 52%;
}

.skeleton div:last-child {
  width: 36%;
}

.people-layout {
  display: grid;
  grid-template-columns: minmax(0, 1.05fr) minmax(0, 0.95fr);
  gap: 18px;
  max-width: var(--max);
  margin: 0 auto;
}

.pi-card {
  display: grid;
  grid-template-columns: 160px minmax(0, 1fr);
  gap: 30px;
  padding: 28px;
  background:
    linear-gradient(135deg, rgba(47, 107, 88, 0.1), rgba(49, 92, 133, 0.1)),
    var(--paper);
}

.initials {
  display: grid;
  place-items: center;
  width: 84px;
  height: 84px;
  border-radius: 8px;
  color: var(--paper);
  background: linear-gradient(135deg, var(--green), var(--burgundy));
  font-size: 1.4rem;
  font-weight: 800;
}

.person-photo-frame {
  position: relative;
  display: inline-grid;
  place-items: center;
  width: 84px;
  height: 84px;
  border-radius: 8px;
  isolation: isolate;
}

.person-photo-frame::before {
  content: "";
  position: absolute;
  inset: -3px;
  z-index: -2;
  border-radius: inherit;
  background: linear-gradient(150deg, rgba(47, 107, 88, 0.9), rgba(49, 92, 133, 0.56), rgba(122, 62, 74, 0.72));
}

.person-photo-frame::after {
  content: "";
  position: absolute;
  inset: -14px;
  z-index: -3;
  border-radius: inherit;
  background: radial-gradient(circle at 50% 18%, rgba(255, 255, 255, 0.72), rgba(47, 107, 88, 0.16) 52%, transparent 72%);
  filter: blur(4px);
}

.person-photo {
  display: block;
  width: 100%;
  height: 100%;
  border: 3px solid var(--paper);
  border-radius: inherit;
  object-fit: cover;
  object-position: center 24%;
  background: var(--wash);
  box-shadow:
    inset 0 0 20px rgba(20, 34, 30, 0.12),
    0 16px 34px rgba(20, 34, 30, 0.16);
}

.pi-card .person-photo-frame {
  width: 150px;
  height: 190px;
  border-radius: 76px 76px 34px 34px;
}

.pi-card .initials {
  width: 120px;
  height: 120px;
}

.role {
  margin: 0 0 6px;
  color: var(--green-2);
  font-size: 0.8rem;
  font-weight: 800;
  text-transform: uppercase;
}

.profile-subheading {
  margin: 22px 0 10px;
  color: var(--ink);
  font-size: 0.82rem;
  font-weight: 800;
  text-transform: uppercase;
}

.experience-disclosure {
  margin-top: 22px;
  border: 1px solid rgba(47, 107, 88, 0.18);
  border-radius: 8px;
  background: rgba(255, 255, 255, 0.5);
}

.experience-disclosure summary {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 12px;
  min-height: 46px;
  padding: 0 14px;
  color: var(--ink);
  cursor: pointer;
  font-size: 0.82rem;
  font-weight: 800;
  text-transform: uppercase;
  list-style: none;
}

.experience-disclosure summary::-webkit-details-marker {
  display: none;
}

.experience-disclosure summary svg {
  width: 18px;
  height: 18px;
  color: var(--green-2);
  transition: transform 180ms ease;
}

.experience-disclosure[open] summary {
  border-bottom: 1px solid rgba(47, 107, 88, 0.16);
}

.experience-disclosure[open] summary svg {
  transform: rotate(180deg);
}

.mini-cv {
  display: grid;
  gap: 10px;
  margin: 20px 0 0;
}

.mini-cv div {
  display: grid;
  grid-template-columns: 70px minmax(0, 1fr);
  gap: 12px;
}

.mini-cv dt {
  color: var(--burgundy);
  font-weight: 800;
}

.mini-cv dd {
  margin: 0;
  color: var(--muted);
}

.experience-list {
  display: grid;
  gap: 14px;
  margin: 0;
  padding: 16px 16px 18px;
  list-style: none;
}

.experience-list li {
  position: relative;
  display: grid;
  gap: 2px;
  padding-left: 18px;
}

.experience-list li::before {
  content: "";
  position: absolute;
  top: 8px;
  left: 0;
  width: 7px;
  height: 7px;
  border-radius: 50%;
  background: var(--green-2);
}

.experience-list span {
  color: var(--burgundy);
  font-size: 0.88rem;
  font-weight: 800;
}

.experience-list strong {
  color: var(--ink);
}

.experience-list em {
  color: var(--muted);
  font-style: normal;
}

.team-notes {
  display: grid;
  gap: 12px;
}

.team-notes article {
  padding: 22px;
}

.team-notes h3 {
  margin-top: 0;
}

.member-grid {
  display: grid;
  grid-template-columns: repeat(3, minmax(0, 1fr));
  gap: 16px;
  max-width: var(--max);
  margin: 18px auto 0;
}

.member-grid[hidden] {
  display: none;
}

.member-card {
  display: grid;
  grid-template-columns: 84px minmax(0, 1fr);
  gap: 18px;
  padding: 22px;
}

.member-card h3 {
  margin-top: 0;
}

.member-card p + p {
  margin-top: 8px;
}

.contact-section {
  color: var(--paper);
  background: var(--green);
}

.contact-inner {
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: 28px;
  max-width: var(--max);
  margin: 0 auto;
}

.contact-inner .section-kicker {
  color: #e4c966;
}

.contact-inner p {
  margin: 18px 0 0;
  color: rgba(255, 255, 255, 0.82);
}

.contact-link {
  min-width: 260px;
  padding: 0 18px;
  border: 1px solid rgba(255, 255, 255, 0.25);
  background: rgba(255, 255, 255, 0.1);
}

.site-footer {
  display: flex;
  justify-content: space-between;
  gap: 16px;
  padding: 28px max(22px, calc((100vw - var(--max)) / 2));
  color: #dbe7e2;
  background: #0d1d18;
  font-size: 0.9rem;
}

.site-footer p {
  margin: 0;
}

@media (max-width: 900px) {
  .site-header {
    padding-inline: 18px;
  }

  .nav-toggle {
    display: grid;
  }

  .site-nav {
    position: fixed;
    inset: 72px 14px auto 14px;
    display: none;
    flex-direction: column;
    align-items: stretch;
    padding: 10px;
    border: 1px solid var(--line);
    border-radius: 8px;
    background: var(--paper);
    box-shadow: var(--shadow);
  }

  .site-nav.is-open {
    display: flex;
  }

  .site-nav a {
    padding: 13px 12px;
  }

  .hero {
    min-height: 78vh;
    padding: 86px 22px 66px;
    background:
      linear-gradient(90deg, rgba(15, 37, 32, 0.96) 0%, rgba(15, 37, 32, 0.72) 70%, rgba(15, 37, 32, 0.38) 100%),
      url("assets/hero-heor-lab.png") center right / cover no-repeat;
  }

  .hero h1 {
    font-size: 2.8rem;
  }

  .hero-lede {
    font-size: 1.08rem;
  }

  .intro-section,
  .research-section,
  .publications-section,
  .people-section,
  .contact-section {
    padding: 66px 22px;
  }

  .section-grid,
  .people-layout,
  .member-grid {
    grid-template-columns: 1fr;
    gap: 28px;
  }

  .section-heading,
  .contact-inner {
    display: grid;
    gap: 18px;
  }

  .section-grid h2,
  .section-heading h2,
  .contact-inner h2 {
    font-size: 2.1rem;
  }

  .research-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .publication-actions {
    justify-content: flex-start;
  }

  .publication-toolbar {
    align-items: flex-start;
    flex-direction: column;
  }

  .publication-year-group {
    grid-template-columns: 1fr;
    gap: 10px;
  }

  .publication-year {
    position: static;
    justify-self: start;
    min-width: 82px;
  }
}

@media (max-width: 620px) {
  .brand strong {
    font-size: 0.84rem;
  }

  .brand small {
    display: none;
  }

  .brand-logo {
    width: 66px;
    height: 38px;
  }

  .brand-mark {
    width: 38px;
    height: 38px;
  }

  .hero h1 {
    font-size: 2.28rem;
  }

  .hero-actions,
  .publication-actions {
    align-items: stretch;
    flex-direction: column;
  }

  .button,
  .contact-link {
    width: 100%;
  }

  .research-grid {
    grid-template-columns: 1fr;
  }

  .research-card {
    min-height: auto;
  }

  .publication-meta {
    display: grid;
  }

  .publication-toolbar {
    gap: 10px;
  }

  .publication-tabs {
    width: 100%;
  }

  .publication-tab {
    min-height: 42px;
    padding: 0 10px;
    font-size: 0.82rem;
  }

  .publication-year-items {
    padding-left: 12px;
  }

  .publication-row {
    grid-template-columns: 1fr;
    gap: 10px;
    padding: 16px;
  }

  .publication-row::before {
    left: -17px;
  }

  .publication-doi-chip {
    justify-self: start;
  }

  .pi-card {
    grid-template-columns: 1fr;
  }

  .member-card {
    grid-template-columns: 1fr;
  }

  .mini-cv div {
    grid-template-columns: 1fr;
    gap: 2px;
  }

  .site-footer {
    display: grid;
  }
}